The 2 theories give some other membership regarding anaphase separa- tion
The brand new Family relations away from Mitosis and you can Meiosis (centered on Darlington). Keep in mind that the fresh new chromosomes was twice in the the first physical appearance when you look at the mitosis, but not about basic (leptotene) phase out of meiotic prophase. It pair within the zygotene, beginning to feel twice (split) from inside the pachytene, consequently they are double over the last level from meiotic prophase (diplotene and you may diakinesis). For the mitosis anaphase initiate by office of the centromeres, however in meiosis it doesn’t are present before the anaphase of the following division (not shown). Darlington are the first to ever advise that if the parasynaptic interpre- tation is used, all the differences between meiosis and you will mitosis is also, with few hypotheses, getting deduced about new difference between the brand new singleness or doubleness of your prophase posts.
The guy ideal you to definitely homologous chromosomes, or rather homologous chromonemata, focus both into the sets. This destination was met when you look at the mitotic prophase, although not in early meiotic prophase, where in fact the chromosomes has reached earliest unmarried, and can simply satisfy the appeal from the future with her inside zygotene combining. Inside meiosis, the latest spHtting of the chromosomes, hence happens in the new interphase just before a good mitotic division, will not occurs right up until pachytene if chromosomes already are associated for the pairs. It thus provides a collection of five associated threads and they break apart to your diplotene loops, becoming kept along with her by alter off spouse from the chiasmata. The newest falling aside of the four posts is proof you to even if a couple of homologous chromosomes notice each other, one collection of homologous chromonemata repels another similar partners.
Hence Darlington supposed that the spUtting from a good chromosome in preparation for another mitotic division happens in the interphase in advance of that section, and this the fresh new singleness of your meiotic chromo- somes is due just to the fact they start condensation and you will con- traction to your section through to the busting has actually took place. So it hypo- thesis is known as the fresh Precocity Principle. A modification of the simple precocity concept could have been recommended by Huskins,^ who accepts Darhngton’s hypotheses your differences between both kinds off department are due to the newest repulsion between sets out-of chromonemata and you may attraction between single chromonemata, however, denies Darlington’s account of resource of your own singleness of the fresh meiotic chromosomes during the prophase. Centered on Huskins, this new splitting of chromosomes for example department occurs throughout the last section, developing a therefore-entitled tertiary broke up that creates the brand new chromo- certain sets on mitotic metaphase are actually quadripartite, people at meiotic metaphase octopartite authorities.
The fresh new Mechanics Of your CHROMOSOMES II5 facts to get telosynapsis try adduced from the density regarding continuous rings out of chromosomes in the meiotic metaphase in a few bacteria (mainly Oenothera), however, another so much more sufficient explana- tion can now get for this (p
Into the interphase prior to meiosis, this tertiary spUt have to be supposed to be destroyed in certain way, merely to be restored during the pachytene; within brand new diakinesis phase it ^ Cf. Huskins and you can Smith 1934. Il6 An overview of Progressive Genetics are been successful from the several other split on the section and therefore observe the second meiotic department. Predicated on Darlington, the first break up of the chromo- somes for the anaphase is actually mitosis as a result of the section off the newest centromeres at this https://datingranking.net/escort-directory/indianapolis point and also in meiosis so you’re able to a heightened repulsion within centromeres and that today starts to beat the pushes holding brand new chromosomes along with her at the chiasmata. Considering Huskins, likewise, the new break up of one’s mitotic chromosomes was analogous toward formation out of diplotene loops ; it is due to the synthesis of the latest tertiary split and that turns new coordinated mitotic chromosomes into the a couple of collectively repelling coordinated threads.
